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Clockwise from top left: Amanita muscaria , a basidiomycete; Sarcoscypha coccinea , an ascomycete; bread covered in mold ; a chytrid; an Aspergillus conidiophore. The evolution of fungi has been going on since fungi diverged from other life around 1. Since fungi do not biomineralise , they do not readily enter the fossil record; there are only three claims of early fungi.

One from the Ordovician [2] has been dismissed on the grounds that it lacks any distinctly fungal features, and is held by many to be contamination; [3] the position of a “probable” Proterozoic fungus is still not established, [3] and it may represent a stem group fungus. There is also a case for a fungal affinity for the enigmatic microfossil Ornatifilum. Since the fungi form a sister group to the animals, the two lineages must have diverged before the first animal lineages, which are known from fossils as early as the Ediacaran.

Factors that likely contribute to the under-representation of fungal species among fossils include the nature of fungal fruiting bodies , which are soft, fleshy, and easily degradable tissues and the microscopic dimensions of most fungal structures, which therefore are not readily evident. Fungal fossils are difficult to distinguish from those of other microbes, and are most easily identified when they resemble extant fungi.

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It is considered as one of the first major Neoglacial advance in the western Alps, in agreement with other regional paleoclimatological proxies. Other Neoglacial advances dated here match the timing of previously described Alpine Neoglacial events. Our results also suggest that a Neoglacial maximum occurred at Etages Glacier 0.

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